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University of the Pacific

Studies on gametogenesis in the Echiuroid worm, Urechis caupo

Abstract

dc:description.abstract

<p>During the spring of 1955, while attending classes at the Pacific Marine Station, under the direction of Dr. Alden E. Nobel, Chairman of the Department of Zoology, College of the Pacific, Urechis caupo was introduced as a representative of the fauna of the mud flats of Tomales Bay. The unusual manner in which this animal lives would stimulate the curiosity of any student interested in biology, and especially those associated with marine life. The fact that the animal burrows its own home in a U-shaped tunnel and dwells there with three permanent commensals (Hesperonoë adventor, Skogsberg, a plynoid annelid; and 2 pinnotherid crabs, Scleroplax granulate, Rathburn, and Pinnixa franciscapa, Rathburn) is a unique characteristic. The goby fish, Clevelandia ios (Jordan and GIlbert), may also be considered a commensal, but it uses the burrow as a retreat rather than a permanent residence (Fisher, 1946).</p><p>Studies on the early embryology of Urechis stimulated the writer to further study of the reproductive cycle. The puzzling descriptions and information pertaining to the immature gametes that are found in the coelomic fluid, coupled with the fact that no permanent gonad has been observed or indeed, that it may be wholly lacking, supplied a challenge that he could not ignore. The primary purpose of this investigation is to attempt to evaluate the reproductive cycle and to attempt to locate some type of germinal epithelium.</p>

Degree

thesis:*
Name thesis:degree_name
Master of Arts (M.A.)
Level thesis:degree_level
Thesis
Discipline thesis:degree_discipline
Zoology
Year dc:date.available
1957

Author and committee

dc:creator, dc:contributor.*
Author dc:creator
  • Hanson, James Charles
Contributors dc:contributor
  • Alden E. Noble

Subjects

dc:subject × 5

Rights

dc:rights

Identifiers

dc:identifier.*
Repository record dc:identifier
https://scholarlycommons.pacific.edu/uop_etds/1335
OAI identifier oai:identifier
oai:scholarlycommons.pacific.edu:uop_etds-2334

Chain of custody

source
Harvested from
University of the Pacific
Base URL
scholarlycommons.pacific.edu/do/oai/
Last updated
2026-07-24
Source record
OAI-PMH GetRecord
citation

Hanson, James Charles. Studies on gametogenesis in the Echiuroid worm, Urechis caupo. Thesis thesis, 1957. https://scholarlycommons.pacific.edu/uop_etds/1335