Diversity, phylogeny and biostratigraphy of diprotodontoids (marsupialia: diprotodontidae, palorchestidae) from the Riversleigh world heritage area

The extinct diprotodontoids were large bodied, browsing herbivorous marsupials most closely related to, among living marsupials, wombats. Referred to two families, Diprotodontidae and Palorchestidae, diprotodontoids are geographically and temporally widespread vombatimorphian taxa in Australian and New Guinean Cenozoic deposits. The most diverse diprotodontoid fauna recorded from any single region in Australia comes from Oligo-Miocene limestone deposits of the Riversleigh World Heritage Area, northwestern Queensland. In this thesis a new diprotodontoid genus and five new species are described from Riversleigh, as well as additional material from Riversleigh for the palorchestids Propalorchestes novaculacephalus and Pr. ponticulus and diprotodontids Nimbadon lavarackorum, Neohelos tirarensis, Neohelos stirtoni and Ngapakaldia bonythoni. A new vombatomorphian family, Maradidae, known from a single species at Riversleigh, is recognised as the sister-group of a vombatid-diprotodontoid clade. New abundant, exceptionally well-preserved cranial material of the zygomaturine Nimbadon lavarackorum enables characterisation of intraspecific variation and ontogenetic development. The results of these analyses have been used to discriminate species boundaries throughout this work. Consequently: Nimbadon whitelawi is now considered a junior synonym of Ni. lavarackorum; Nimbadon scottorrorum is a junior synonym of Neohelos tirarensis; and Bematherium angulum is a synonym of Ngapakaldia bonythoni. The new Riversleigh diprotodontoids clarify phylogenetic relationships within and between diprotodontoid families. The monophyly of both Palorchestidae and Diprotodontidae is strongly supported as is their union in the superfamily Diprotodontoidea. Monophyly of the Zygomaturinae and Diprotodontinae is not supported, primarily due to the unstable position of Alkwertatherium webbi as well as the high degree of homoplasy in cranial morphology of the more derived members of each subfamily. Overall phylogenetic and distribution patterns for diprotodontoids is generally consistent with current interpretations of Riversleigh's stratigraphy. Five diprotodontoid species allow direct biocorrelation with other Australian Tertiary mammal faunas. Riversleigh's basal System A deposits correlate with late Oligocene deposits of the Etadunna Formation of South Australia. Riversleigh's low-mid System C deposits correlate with the middle Miocene Bullock Creek Local Fauna of the Northern Territory. Riversleigh's high System C Jaw Junction and Encore Local Faunas contain diprotodontoid taxa antecedent to diprotodontoids of the late Miocene Alcoota Local Fauna of the Northern Territory.

Grantor dc:publisher

UNSW, Sydney

Year dc:date

2008

Author dc:creator

• Black, Karen

• Systematics.
• Diprotodontoidea.
• Ontogeny.
• Maradidae.
• Oligocene.

Statement dc:rights

• open access
• CC BY-NC-ND 3.0
• free_to_read

Licence

• https://purl.org/coar/access_right/c_abf2
• https://creativecommons.org/licenses/by-nc-nd/3.0/au/

Language dc:language

EN

Identifier

https://doi.org/10.26190/unsworks/2719

OAI identifier oai:identifier

oai:unsworks.library.unsw.edu.au:1959.4/43327