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Loma Linda University

Modern Invertebrate Footprints Compared with Fossil Footprints from the Coconino Sandstone (Permian)

Abstract

dc:description.abstract

<p>Laboratory studies of trackways of the crayfish, Cambarus clarkii were conducted, and the footprints compared with fossil invertebrate footprints (Paleohelcura sp.) of the Permian Coconino sandstone of Northern Arizona, and similar footprints of other sandstone. Fine grained sand (similar to that composing the Coconino sandstone) was placed in a plexiglass tank to form slopes of various angles. The crayfish walked up and down slopes of 10°± 3°, 20°± 2°, 25° and on level sand, and their trackways were photographed. Trackways were produced on wet sand and underwater sand, but the animals would not walk on dry sand. Clear individual impressions (footprints) were present in one hundred percent of the wet sand trackways and ninety percent of the underwater trackways. A variety of track patterns was produced with twenty percent of the underwater trackways resembling the fossil track patterns of Paleohelcura sp. Forty-five percent of all the underwater trackways and eighty-eight percent of those made on low dipping slopes we re also very similar to Paleohelcura sp. in characteristic features of the individual impressions. These data indicate that trackways such as those named Paleohe lcura could have been made in an underwater environment by some aquatic animal, such as a crustacean.</p>

Degree

thesis:*
Name thesis:degree_name
Master of Arts (MA)
Level thesis:degree_level
Thesis
Discipline thesis:degree_discipline
Biology
Year
1977

Author and committee

dc:creator, dc:contributor.*
Author dc:creator
  • Emtage, Marilyn A.
Contributors dc:contributor
  • Leonard R. Brand
  • Arthur E. Dalgleish
  • Conrad D. Clausen

Subjects

dc:subject × 2

Rights

dc:rights
Statement dc:rights
  • This title appears here courtesy of the author, who has granted Loma Linda University a limited, non-exclusive right to make this publication available to the public. The author retains all other copyrights.
Language dc:language
English

Identifiers

dc:identifier.*
Repository record dc:identifier
https://scholarsrepository.llu.edu/etd/590
OAI identifier oai:identifier
oai:scholarsrepository.llu.edu:etd-1723

Chain of custody

source
Harvested from
Loma Linda University
Base URL
scholarsrepository.llu.edu/do/oai/
Last updated
2026-07-24
Source record
OAI-PMH GetRecord
citation

Emtage, Marilyn A.. Modern Invertebrate Footprints Compared with Fossil Footprints from the Coconino Sandstone (Permian). Thesis thesis, 1977. https://scholarsrepository.llu.edu/etd/590