Male genital accessory gland development in Dermacentor variabilis is completed by the fourth day after initiation of feeding, while that in Argas radiatus (Argasidae) occurs between the first and tenth days after adult ecdysis. D. variabilis has twelve spongy and/or granular lobes, and A. radiatus has fourteen spongy and/or granular lobes. The glands lack cellular differentiation in immatures, but have four distinct cell types present in adults. Both species also possess apocrine, holocrine, and merocrine secretory mechanisms. Secretions contain acid polysaccharides, mucin, glycogen, basic protein, and other products. Continually produced accessory gland secretions accumulate before mating and are discharged at the time of mating. Additional secretory material is produced soon after mating. A new tubular cell type is described.
","abstract_html":"<p>Male genital accessory gland development in <em>Dermacentor</em> <em>variabilis</em> is completed by the fourth day after initiation of feeding, while that in <em>Argas</em> <em>radiatus</em> (Argasidae) occurs between the first and tenth days after adult ecdysis. <em>D</em>. <em>variabilis</em> has twelve spongy and/or granular lobes, and A. radiatus has fourteen spongy and/or granular lobes. The glands lack cellular differentiation in immatures, but have four distinct cell types present in adults. Both species also possess apocrine, holocrine, and merocrine secretory mechanisms. Secretions contain acid polysaccharides, mucin, glycogen, basic protein, and other products. Continually produced accessory gland secretions accumulate before mating and are discharged at the time of mating. Additional secretory material is produced soon after mating. A new tubular cell type is described.</p>","abstract_has_math":false,"creators":["Murphy, Richard W."],"institution":null,"degree_name":"Master of Science in Biology","degree_level":"Thesis (restricted to Georgia Southern)","degree_discipline":"Department of Biology","degree_department":null,"school":null,"contributors":["W. Keith Hartberg","Edwin T. Hibbs"],"advisors":[],"committee_chairs":[],"committee_members":[],"year":1974,"date_issued":"1974-01-01T08:00:00Z","date_published":"1974-01-01T08:00:00Z","updated_at":"2026-07-24T02:28:40Z","subjects":["ETD","Male genital accessory gland","Dermacentor variabilis","Argas radiatus","Secretions","Histology","Histochemistry","Biochemistry, Biophysics, and Structural Biology","Biology"],"languages":[],"rights":[],"rights_urls":[],"identifier_entries":[]},"links":{"outbound_url":"https://digitalcommons.georgiasouthern.edu/etd_legacy/821","outbound_label":"Repository record","outbound_source":"dc:identifier"},"metadata_groups":[{"id":"people","label":"People","entries":[{"key":"dc:contributor","label":"Contributor","values":["W. Keith Hartberg","Edwin T. Hibbs"]},{"key":"dc:creator","label":"Author","values":["Murphy, Richard W."]}]},{"id":"academic_context","label":"Academic Context","entries":[{"key":"dc:date.available","label":"Dc Date Available","values":["2015-09-15T07:00:00Z"]},{"key":"thesis:degree_discipline","label":"Discipline","values":["Department of Biology"]},{"key":"thesis:degree_level","label":"Degree Level","values":["Thesis (restricted to Georgia Southern)"]},{"key":"thesis:degree_name","label":"Degree Name","values":["Master of Science in Biology"]}]},{"id":"subjects_keywords","label":"Subjects and Keywords","entries":[{"key":"dc:subject","label":"Dc Subject","values":["ETD","Male genital accessory gland","Dermacentor variabilis","Argas radiatus","Secretions","Histology","Histochemistry","Biochemistry, Biophysics, and Structural Biology","Biology"]}]},{"id":"identifiers","label":"Identifiers","entries":[{"key":"dc:identifier","label":"Identifier","values":["https://digitalcommons.georgiasouthern.edu/etd_legacy/821"]}]},{"id":"additional","label":"Additional Metadata","entries":[{"key":"dc:description.abstract","label":"Abstract","values":["Male genital accessory gland development in Dermacentor variabilis is completed by the fourth day after initiation of feeding, while that in Argas radiatus (Argasidae) occurs between the first and tenth days after adult ecdysis. D. variabilis has twelve spongy and/or granular lobes, and A. radiatus has fourteen spongy and/or granular lobes. The glands lack cellular differentiation in immatures, but have four distinct cell types present in adults. Both species also possess apocrine, holocrine, and merocrine secretory mechanisms. Secretions contain acid polysaccharides, mucin, glycogen, basic protein, and other products. Continually produced accessory gland secretions accumulate before mating and are discharged at the time of mating. Additional secretory material is produced soon after mating. A new tubular cell type is described.
"]},{"key":"dc:title","label":"Title","values":["Comparative Histology and Histochemistry of the Male Genital Accessory Glands of Dermacentor variabilis (Say and Argas radiatus (Railliet)"]}]}],"canonical_facts":{"dc:contributor":["W. Keith Hartberg","Edwin T. Hibbs"],"dc:creator":["Murphy, Richard W."],"dc:date.available":["2015-09-15T07:00:00Z"],"dc:description.abstract":["Male genital accessory gland development in Dermacentor variabilis is completed by the fourth day after initiation of feeding, while that in Argas radiatus (Argasidae) occurs between the first and tenth days after adult ecdysis. D. variabilis has twelve spongy and/or granular lobes, and A. radiatus has fourteen spongy and/or granular lobes. The glands lack cellular differentiation in immatures, but have four distinct cell types present in adults. Both species also possess apocrine, holocrine, and merocrine secretory mechanisms. Secretions contain acid polysaccharides, mucin, glycogen, basic protein, and other products. Continually produced accessory gland secretions accumulate before mating and are discharged at the time of mating. Additional secretory material is produced soon after mating. A new tubular cell type is described.
"],"dc:identifier":["https://digitalcommons.georgiasouthern.edu/etd_legacy/821"],"dc:subject":["ETD","Male genital accessory gland","Dermacentor variabilis","Argas radiatus","Secretions","Histology","Histochemistry","Biochemistry, Biophysics, and Structural Biology","Biology"],"dc:title":["Comparative Histology and Histochemistry of the Male Genital Accessory Glands of Dermacentor variabilis (Say and Argas radiatus (Railliet)"],"thesis:degree_discipline":["Department of Biology"],"thesis:degree_level":["Thesis (restricted to Georgia Southern)"],"thesis:degree_name":["Master of Science in Biology"]},"updated_at":"2026-07-24T02:28:40Z"}