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East Tennessee State University

Systematics, Climate, and Ecology of Fossil and Extant Nyssa (Nyssaceae, Cornales) and Implications of Nyssa grayensis sp. nov. from the Gray Fossil Site, Northeast Tennessee

Abstract

dc:description.abstract

<p>The Late Hemphillian (latest Miocene or earliest Pliocene, 7-4.5 Ma) Gray Fossil Site in northeastern Tennessee is interpreted to represent a lacustrine paleokarst fed by a river or stream. This research focuses on the morphological and systematic relations of Nyssa endocarps (fruit pits) from the fossil site to extinct and extant Nyssa species. A combination of metric and nonmetric traits allows recognition of a new species: <em>Nyssa grayensis</em> sp. nov. This fossil species shares the most similarities with the extant <em>Nyssa ogeche</em> Bartram ex Marshall from southeast North America and the Eocene fossil <em>Nyssa eolignitica</em> Berry from western Tennessee. Affinities with <em>Nyssa ogeche</em> Bartram ex Marshall suggest a warmer winter climate than the present and periodically fluctuating water levels. Fossil vertebrates (<em>Alligator, Heloderma, Hesperotestudo</em>) and plants (Fossil relatives of <em>Ilex vomitoria, Quercus virginiana, Taxodium, Acer leucoderme</em>) with modern counterparts distributed in areas with mild winters support this interpretation.</p>

Degree

thesis:*
Name thesis:degree_name
MS (Master of Science)
Level thesis:degree_level
Thesis - unrestricted
Discipline thesis:degree_discipline
Biology
Year dc:date.issued
2013

Author and committee

dc:creator, dc:contributor.*
Author dc:creator
  • Noll, Nathan R

Subjects

dc:subject × 10

Rights

dc:rights
Statement dc:rights
  • Copyright by the authors.

Identifiers

dc:identifier.*
Repository record dc:identifier
https://dc.etsu.edu/etd/1204
OAI identifier oai:identifier
oai:dc.etsu.edu:etd-2366

Chain of custody

source
Harvested from
East Tennessee State University
Base URL
dc.etsu.edu/do/oai/
Last updated
2026-07-24
Source record
OAI-PMH GetRecord
citation

Noll, Nathan R. Systematics, Climate, and Ecology of Fossil and Extant Nyssa (Nyssaceae, Cornales) and Implications of Nyssa grayensis sp. nov. from the Gray Fossil Site, Northeast Tennessee. Thesis - unrestricted thesis, 2013. https://dc.etsu.edu/etd/1204